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  1. Beck, D.D., 2005
    Biology of Gila Monsters and Beaded Lizards.
    University of California Press Ltd., London, England

  2. Bogert, C.M. and R. Martin del Campo, 1956
    The Gila Monster and its Allies.
    The relationships, habits, and behavior of the lizards of the family Helodermatidae.
    Bulletin of the American Museum of Natural History 109: 1-238.

  3. Scull of Heloderma susp. out of the collection of the Senckenberg Institute Frankfurt/Main, Germany.
    Curtesy of Dr. G. Köhler to take photographs.

  4. Russel. F.E., and C.M. Bogert, 1981
    Gila-Monster. Its biology, venom and bite - A review.
    Toxicon 19: 341-59.

  5. Beck, D.D., 2005
    Biology of Gila Monsters and Beaded Lizards.
    The University of California Press, License No. 2106511339325

  6. Nydam, R.L., 2000.
    A new taxon of helodermatid-like lizard from the Albian-Cenomanian of Utah.
    Journal of Vertebrate Paleontology 20 (2): 285-94.

  7. Mead, J.I., E.L. Roth, T.R. Van Devender, and D.W. Steadman, 1984
    The late Wisconsinan vertebrate fauna from Deadman Cave, southern Arizona.
    Transactions of the San Diego Society of Natural Hiostory 20 (14): 247-76.

  8. Beck, D.D., 1990
    Ecology and behavior of the Gila-Monster in southwestern Utah.
    Journal of Herpertology 24 (I): 54-68.

  9. Cooper, W.E., Jr. and I. Arnett, 1995
    Strike-induced chemosensory searching in the Gila Monster.
    Copeia 1995 (I): 89-96.

  10. Wegscheider, F.J., 1998
    Assimilation efficiency of the Gila Monster Heloderma suspectum.
    MS thesis. Fullerton: California State University.

  11. DeNardo, Dale F., Zubal, Tricia E. and Hoffman, Ty C. M., 2004
    Cloacal evaporative cooling: a previously undescribed means of increasing evaporative water loss at higher temperatures in a desert ectotherm, the Gila monster Heloderma suspectum.
    The Journal of Experimental Biology 207, 945-953

  12. Beck, D.D., and R.D. Jennings, 2003
    Habitat use by Gila Monsters. The importance of shelters.
    Herpetological Monographs 17: 112-130.

  13. Lowe, C.H., P.J. Lardner and E.A. Halpern, 1971
    Supercooling in reptiles and other vertebrates.
    Comparative Biochemistry and Physiology 39A: 125-135.

  14. Mattlin, A.K., L.I. Gallardo and D.F. DeNardo, 2003
    Monsters on the move: Activity patterns in Heloderma suspectum.
    Unpublished poster presentation, Department of Biologie. Tempe: Arizona State University.

  15. Gienger, C.M., 2003
    Natural history of the Gila Monster in Nevada.
    MS thesis. Reno: University of Nevada.

  16. Brown, D.E. and N.B. Carmoney. 1999.
    Facts and Folklore of America´s Aztec lizard.
    The University of Utah Press, 1999

  17. Beck, D.D., 2004
    Venomous lizards of the desert.
    Natural History, July/August 32-36

  18. Douglas, M.E., M.R. Douglas, G.W. Schuett, D.D. Beck, and B.K. Sullivan. 2003.
    Molecular biodiversity of Helodermatidae
    (Reptilia, Squamata). Abstract and presentation.
    Joint Meeting of Ichthyologists and Herpetologists. Manaus, Amazonas, Brazil.

  19. Lit.1, p.132-134

  20. Lit.1, p.149

  21. Lowe, C.H., C.R. Schwalbe, and T.B. Johnson, 1986
    The Venomous Reptiles of Arizona.
    Phoenix: Arizona Game and Fish Department.

  22. Goldberg, S.R., and C.H. Lowe, 1997
    Reproductive cycle of the Gila Monster, Heloderma suspectum, in southern Arizona.
    Journal of Herpetology 31:161-66

  23. Greek, T.J., 1991
    Operative temperature for the herpetoculturist, 1991 Capative Progation and Husbandary Conference,
    1991 Davis Norhern Herpetological Society Special Publication.

  24. Loza, E. 2001. Natural history and captive husbandary of the Gila Monster.
    The Reptile and Amphibian Hobbyist. Vol 6 (5) 10-19.

  25. Seward, M., 1998
    Captive reproduction of the Gila Monster
    Reptiles 6(12) 68-75

  26. Seward, M., 2002
    Dr. Marc Seward´s Gila Monster Propagation: How to bread Gila Monsters in captivity.
    Colorado Springs, CO. : Natural Selections Publishing.

  27. Peterson, K.H., 1982
    Reproduction in captive Heloderma suspectum.
    Herpetological Review13(4)122-124.

  28. Eidenmüller, B. und Wicker. R., 1992
    Über eine Nachzucht von Heloderma suspectum.
    Salamandra 28/2 106-111

  29. Eidenmüller, B. 1992. Bemerkungen zur Nachzucht von Heloderma suspectum (Cope, 1869).
    Monitor 1(1) 24-32

  30. Steinfartz, S., 1993
    Über eine Nachzucht von Heloderma suspectum (Cope, 1869) im Terrarium.
    Sauria 15(1) 3-8

  31. Demeter, J.B. 1986
    Combat behavior in the Gila Monster (Heloderma suspectum cinctum).
    Herpetological Review, Lawrence 17(1)9-10

  32. DeNardo, D., 1991
    Egg Retention in Reptilles.
    1991 NHCS Captive Propagation and Husbandary Conference, Davis,
    Northern California Herpetological Society (NHCS) Special Publication.

  33. Switak, K., 1984
    The life of desert reptiles and amphibiens.
    Karl. H. Switak Press. P.O. Box 27141, San Francisco, CA 94127, Feb. 1984

  34. Sullivan, Brian K., Kwiatkowski, Matthew A., Schuett, Gordon W., 2004
    Translocation of urban Gila Monsters: a problematic conservation tool
    Biological Conservation 117: 235-242

  35. Morris, Patrick J. and Henderson, Charles, 1998
    Gender Determination in Mature Gila Monster, Heloderma suspectum, and Mexican Beaded Lizards,
    Heloderma horridum, by Ultrasound Imaging of the Ventral Tail.
    Assoc. Reptillian Amphibian Vet, Vol. 8 No. 4

  36. Vidal, N. and Hedges, S. B. 2005
    The phylogeny of squamate reptiles (lizards, snakes, and amphisbaenians) inferred from nine nuclear protein-coding genes
    C. R. Biologies 328 (2005) 1000 - 1008

  37. Lidikay, Charles E. and Stone, K., 1997
    Biochemistry of Helodermatid Venom

  38. Mebs, D "Gifttiere", 2000
    Wissenschaftliche Verlagsges, 2. Auflage, neu bearb., 248-250 (2000)

  39. Odermatt, C. (Buochs), 1940
    Beiträge zur Kenntnis des Gebisses von Heloderma
    Vierteljahresschrift der Naturforschenden  Gesellschaft in Zürich LXXXV

  40. Eidenmüller, B. und M. Reisinger, 2011
    Krustenechsen – Lebensweise, Haltung und Zucht.
    Edition Chimaira  2011

  41. Edmund, A. G., 1969
    Biology of the Reptilia (C. Gans, A. d'A Bellairs and T. S. Parsons, eds.). Vol. 1, pp. 117-200, Academic Press, London

  42. King, G., 1996
    Reptiles and Herbivory 
    London: Chapman and Hall, Lizards , 29- 42

  43. Shaw, C.E., 1968
    Reproduction of the Gila Monster (Heloderma suspectum) at the San Diego Zoo
    Zool. Garten 35(1/2) 1-6, Leipzig

  44. Shufeldt, R.W. - M.D., C.M.Z.S., 1890
    Contributions to the Study of Heloderma suspectum
    Proc. Zool. Soc. of London, April 1, 149-244

  45. Salomon, F.- V., W. Achilles und U. Gille,  2008
    Anatomie für die Tiermedizin
    Enke Verlag Stuttgard 2008

  46. O’Malley, Baibre, 2008
    Klinische Anatomie und Physiologie bei kleinen Heimtieren,Vögeln, Reptilien und Amphibien, Kap. 4- Echsen  p. 70- 89
    Verlag: Elsevier, München: Urban & Fischer 2008

  47. Versluys, J., 1899
    Die mittlere und äussere Ohrsphäre der Lacertilia und Rhynchocephalia
    Fam. Helodermatidae  20. Heloderma suspectum Cope
    Zool. Jahrbücher, Band XII,  219- 223
    Verlag Gustav Fischer, Jena 1899

  48. Schmidt, J. W., 1912
    Studien am Integument der Reptilien
    II Die Hautverknöcherungen von Heloderma
    Zool. Jahrbücher, Suppl. 15, II, 219- 228
    Verlag Gustav Fischer, Jena 1912

  49. Bellairs, A., 1969
    The Life of Reptiles  Sex and Reproduction
    London: Weidenfeld and Nicolson II 390 -433

    Bellairs, A.
    The Life of Reptiles - Die Reptilien
    Wermuth, H., Bearbeitung und Übersetzung ins Deutsche 1971
    Embryonal-Entwicklung, Chromosomen und Parthenogenese
    Band 11, Kapitel 9 (517 - 537)
    Edition Rencontre Lausanne 1971

  50. Köhler, G. 1997
    Inkubation von Reptilieneiern: Grundlagen, Anleitungen, Erfahrungen
    Herpeton Verlag, Offenbach 1997

  51. Brehms Thierleben, 1878
    Allgemeine Kunde des Thierreichs, Band 1, 3. Abteilung, Kriechtiere, Lurche und Fische,
    Krustenechsen 182 - 183
    Verlag des Bibliographischen Institutes, Leipzig 1878

  52. Nowak. E. 2005
    Why did the Gila Monster Cross the Road? Results from a Study at Toronto National Monument, Arizona,
    Sonoran Herpetologist 18 (9) 2005

  53. Douglas, Michael E., Marlis R. Douglas, Gordon W. Schäfer, Daniel D. Beck, Brian K. Sullivan, 2010
    Conservation phylogenetics of helodermatid lizards using murliple molecucar markers and supertree approach
    Molecular Phylogenetics and Evolution 55, 153 - 167

  54. Bech, R.,und U. Kaden 1990
    Vermehrung von Terrarientieren - Echsen
    Urania Verlag Leipzig, Jana, Berlin 1990

  55. Beck, D.and „Wild Earth Guardians“ 2010
    Petition to List the Utah Population of the Gila Monster (Heloderma suspectum) under the U. S. Endangered Species Act

  56. Stitt, Eric W., Cecil R. Schwalbe, and Don E. Swann 2003
    Signs of Gila Monster Predation on Desert Tortoise Nests
    Sonoran Herpetologist – Tucson 2003

  57. Davis Jon R. and Dale F. DeNardo, 2007
    The urinary bladder as a physiological reservoir that moderates dehydration in a large desert lizard,
    the Gila monster Heloderma suspectum
    The Journal of Experimental Biology, 210 (1472- 1480)

  58. Judd, Howard L., James P. Bacon, Dieter Rüedi, J. Girard and K. Benirschke, 1977
    Determination of Sex in the Komodo Dragon International Zoo Book

  59. Mayes, P. J., Bradshaw S. D., and Bradshaw, F. J., 2005
    Successfully Determining the Sex of Adult Varanus mertensi (Reptilia: Varanidae) Using a Combination of Both Hemipenile Eversion and the Ratio of Androgens: Estradiol in Plasma
    Ann. N. Y. Acad. Sci. 1040 (402 -405)

  60. Fry, Bryan G., Vida Nicolas, l et al. , 2006
    Early Evolution of the Venom System in Lizards and Snakes Nature 439 (584- 588)

  61. Bhullar, Bhart- Anjan S. and Krister Smith, 2008
    Helodermatid Lizard from the Miocene of Florida, the Evolution of the Dentary in Helodermatidae, and Comments on Dentary Morphology in Varanoidea
    Journal of Herpetology Vol. 2, No. 2, (286- 302)

  62. Kerri Cooper, Bailey, Stephen A. Smith, Kurt Zimmerman, Rosalie Lane, Rose E. Raskin, and Dale DeNardo, 2011
    Hematology, leukocyte cytochemical analysis, plasma biochemistry, and plasma electrophoresis of wild- caught and captive- bred Gila monsters (Heloderma suspectum)
    American Society for Veterinary Clinical Pathology (1- 8)

  63. Koludarow, Ivan, Timothy : W. Jackson, et al., 2014
    Fossilized Venom: The Unusually Conserved Venom Profiles of Heloderma Species (Beaded Lizards and Gila Monsters)
    Toxins 6 (3582- 3595)

  64. Gienger, C. M. and Beck, Daniel D,.2007
    Heads or tails? Sexual dimorphism in helodermatid lizards
    Canadian Journal of Zoology 85 (92- 95)

  65. König, Rudolf, 1983
    Schlangen. "Über Bau und Leben faszinierender Tiere".
    Begleitbuch zur Sonderausstellung: "Schlangen und andere lebende Reptilien".
    Zoologisches Museum der Universität Kiel.

  66. Smith, Krister T., Bhart-Anjan S. Bhullar, G. Köhler, J. Habersetzer, 2018
    The Only Known Jawed Vertibrate with Four Eyes and Bauplan of the Pineal Komplex.
    Current Biology, Vol. 28, 1101-1107.

  67. Repp,Roger A., 2017
    A Monstrous Helloween Treat.
    Bulletin of the Chicago Herpetological Society 52(2), 28-35.

  68. Garman, S., 1890
    "On the Gila Monster" (Hdoderma stupectum),
    [strange spelling in the origin title]
    Bulletin of the Essex Institute.
    Vol. XXII, Nos. 4,5 and 6.

  69. DeNardo DF, Moeller KT, Seward M, Repp R., 2018
    Evedence for atypical nest overwintering by hatchling lizards, Heloderma suspectum.
    Proc. Royal Society B. 285: 20180632.

  70. Repp, Roger A., 2015
    The Cyclops Begs Some Bunnies.
    Bull. Chicago Herp. Soc. (8): 127-131.

  71. Davis, J. R., DeNardo, D. F., 2010
    Seasonal patterns of body condition, hydration state, and activity of Gila monsters (Heloderma suspectum) at a Sonoran Desert Site. .
    J. Herpetol. 44, 83- 93. (doi: 10.1670/08-263.1).

  72. Davis, J. R., DeNardo, D. F., 2009
    Water supplementation affects the behavioral and physiological ecology of Gila monsters (Heloderma suspectum) In the Sonoran Desert.
    Physiol. Biochem. Zool. 82, 739- 748 (doi:10.1086/605933).

  73. Hiller, U. 1998
    The Conjunctiva and Nictitating Membrane of the Eyelids in Some Reptilian Species: a SEM Overview
    European Journal of Morphology 36(3), 165- 171 (1998)

  74. Stahnke ,H. L. , 1952
    A Note on the Food of the Gila Monster, Heloderma suspectum Cope
    Herpetologica 8 (3) , 64-65 1952

  75. Grant, C., 1952
    Probably the First Legislation to Protect a Poisonous Animal
    Herpetologica 8 (3) , 64 ( 1952)

  76. Reisinger, M., 2013
    Außergewöhnliches Ereignis bei der Nachzucht des Gila- Monsters (Heloderma s. suspectum)
    Reptilia 18 (6) 2- 4 (2013)

  77. Davis,J. R. and DeNardo, D.F., 2009
    Water supplementation Affects the Behavioral and Physiological Ecology of Gila Monsters (Heloderma suspectum)
    Physiological and Biochemical Zoology 82 (6), 739- 748 (2009)

  78. Hoser, R. T., 2013
    Not in Heloderma … A revised taxonomy and new genus for the Gila Monster.
    Australian Journal of Herpetology 21, 37- 40 (2013)

  79. Zylstra, E. T., Seward, M. T., Schuett, G. W., Repp, R., DeNardo, D. F., Beck, D. D., 2015
    Heloderma suspectum (Gila Monster), Probable Courtship and Mating Behavior.
    Herpetological Review 46(2), 258-259, 2015

  80. Schaal, S. F. K., Smith, K. T. und Habersetzer, J. 2018
    Messel- ein fossiles Tropenökosystem.
    Messel- An Ancient Greenhouse Ecosystem.
    Schweizerbart`sche Verlagsbuchhandlung, Senckenberg- Buch 79

  81. Girling S. J. & Rati P. 2019
    BSAVA: Manual of Reptiles, 3rd Edition
    British Small Animal Veterinary Association Quedgeley, England 1-480.

  82. Holm J. F., 1879
    Some notes on the histology of the poison glands of Heloderma suspectum
    Anatomischer Anzeiger 13, Seiten 80-85.

  83. Chen, Tianbao, H.K. Kwok, C. Ivanyi, Cristipher Shae, 2006
    Isolation and cloning of exendin precursor cDNAs from single samples of venom from the Mexican beaded lizard (Heloderma horridum) and the Gila monster (Heloderma suspectum). Hrsg.: Toxicon. Band 47, Nr. 3, 2006, S. 288–295.

  84. M. K. K. Yap, N. Misuan, 2019
    Exendin‐4 from Heloderma suspectum venom: From discovery to its latest application as type II diabetes combatant. In: Basic Clin Pharmacol Toxicol. Band 124, 2019, S. 513–527.